Mantis shrimp or stomatopods are marine crustaceans, the members of the order Stomatopoda. They may reach 30 centimetres (12 in) in length, although exceptional cases of up to 38 cm (15 in) have been recorded. The carapace of mantis shrimp covers only the rear part of the head and the first four segments of the thorax. Mantis shrimp appear in a variety of colours, from shades of browns to bright neon colours. Although they are common animals and among the most important predators in many shallow, tropical and sub-tropical marine habitats they are poorly understood as many species spend most of their life tucked away in burrows and holes.
Smashers, on the other hand, possess a much more developed club and a more rudimentary spear (which is nevertheless quite sharp and still used in fights between their own kind); the club is used to bludgeon and smash their meals apart. The inner aspect of the dactyl (the terminal portion of the appendage) can also possess a sharp edge, with which the animal can cut prey while it swims.
Both types strike by rapidly unfolding and swinging their raptorial claws at the prey, and are capable of inflicting serious damage on victims significantly greater in size than themselves. In smashers, these two weapons are employed with blinding quickness, with an acceleration of 10,400 g (102,000 m/s2 or 335,000 ft/s2) and speeds of 23 m/s from a standing start, about the acceleration of a .22 calibre bullet. Because they strike so rapidly, they generate cavitation bubbles between the appendage and the striking surface. The collapse of these cavitation bubbles produces measurable forces on their prey in addition to the instantaneous forces of 1,500 newtons that are caused by the impact of the appendage against the striking surface, which means that the prey is hit twice by a single strike; first by the claw and then by the collapsing cavitation bubbles that immediately follow. Even if the initial strike misses the prey, the resulting shock wave can be enough to kill or stun the prey.
The snap can also produce sonoluminescence from the collapsing bubble. This will produce a very small amount of light and high temperatures in the range of several thousand kelvins within the collapsing bubble, although both the light and high temperatures are too weak and short-lived to be detected without advanced scientific equipment. The light emission and temperature increase probably have no biological significance but are rather side-effects of the rapid snapping motion. Pistol shrimp produce this effect in a very similar manner. Smashers use this ability to attack snails, crabs, molluscs and rock oysters; their blunt clubs enabling them to crack the shells of their prey into pieces. Spearers, on the other hand, prefer the meat of softer animals, like fish, which their barbed claws can more easily slice and snag.